{"schema":"https://ver.cy/schemas/card/1.0.0","id":"vr.tr.klepton","code":"thing-q931051","url":"https://ver.cy/models/thing/q931051/","name":"klepton","alternateNames":[],"kind":"thing","status":"published","version":"0.1.0-research.1","language":"en","classifiers":{"family":"Thing Registry","category":"Physical world and living systems","entryKind":"thing","plane":"PHY","domain":["PHY.LIV"],"industry":[],"navPath":"","tags":[],"facets":{}},"whatItIs":"A klepton is a hybrid-derived group of animals whose reproduction depends on genetic contributions or reproductive assistance from a related species.","purpose":"Enable an agent to recognise a klepton as a biological taxonomic entity characterised by reproductive dependence on another taxon, assess the supporting evidence, and evaluate actions affecting that dependence.","scope":{"in":["Authority-specific meaning and circumscription of a klepton","Dependence on other taxa for reproduction","Hybrid ancestry and transmission of parental genomes","Variation in reproductive systems between populations","Evidence needed to identify and monitor klepton populations"],"out":["Legal identity, membership and governance of organisations","Complete models of the parental or reproductive partner taxa","Individual organism anatomy and clinical condition","Hybridisation and reproductive mechanisms as general processes","Whole ecosystem management beyond the klepton's reproductive dependencies"],"boundaries":[]},"distinguishingFeatures":["Classification requires evidence about reproductive dependence; hybrid ancestry alone does not establish a klepton.","The entity is a taxon or circumscribed group of populations, rather than an individual hybrid specimen.","A persistent reproductive lineage must be distinguished from repeated production of transient first-generation hybrids.","Historical ancestry and present reproductive partners must be recorded separately because they need not represent the same relationship.","Use of 'kl.' in a scientific name is evidence of a nomenclatural treatment, but does not independently demonstrate the reproductive mechanism."],"structure":{"bundles":[{"id":"klepton-identity","name":"Klepton identity","description":"Establish what biological entity the registry name denotes and which taxonomic treatment is being applied.","layers":[{"id":"registry-referent","name":"Registry referent","description":"Resolve the intended meaning before assigning biological attributes.","findings":[{"id":"biological-meaning","name":"Biological meaning","description":"Research uses klepton for hybridogenetic biological entities; this supports a provisional biological interpretation of the otherwise undefined registry label. Source: [Scientific Reports study](https://www.nature.com/articles/s41598-017-12942-y).","questions":[{"text":"Does the source record behind vr.tr.klepton identify the biological concept or a different referent?","id":"biological-meaning-q01","kind":"provenance"},{"text":"Which registry plane and domain should represent the confirmed referent?","id":"biological-meaning-q02","kind":"boundary"}]}]},{"id":"taxonomic-circumscription","name":"Taxonomic circumscription","description":"Keep the designation, biological membership and chosen authority connected.","findings":[{"id":"designation-and-membership","name":"Designation and membership","description":"Record the scientific name, klepton notation where used, authorial treatment and criteria for including populations; do not infer membership from the label alone.","questions":[{"text":"Which authority defines this klepton and what populations or lineages does it include?","id":"designation-and-membership-q01","kind":"definition"},{"text":"Which alternative names or treatments refer to the same circumscribed entity?","id":"designation-and-membership-q02","kind":"boundary"}]}]}]},{"id":"reproductive-dependence","name":"Reproductive dependence","description":"Describe the external biological input needed to maintain the klepton.","layers":[{"id":"partner-roles","name":"Partner roles","description":"Identify contemporary reproductive partners independently of ancestral taxa.","findings":[{"id":"required-partner-input","name":"Required partner input","description":"Represent each demonstrated partner relationship with its contribution, compatibility conditions and population context.","questions":[{"text":"Which taxon supplies the reproductive input, and what input is required?","id":"required-partner-input-q01","kind":"definition"},{"text":"What observations distinguish obligatory dependence from occasional successful mating?","id":"required-partner-input-q02","kind":"provenance"}]}]},{"id":"reproductive-mechanism","name":"Reproductive mechanism","description":"Determine what happens to partner input during reproduction.","findings":[{"id":"input-fate","name":"Input fate","description":"Separate developmental activation from inherited genetic contribution. Gynogenesis and hybridogenesis involve different transmission patterns, so they must not be treated as interchangeable. Source: [Evolutionary perspectives on clonal reproduction in vertebrate animals](https://pmc.ncbi.nlm.nih.gov/articles/PMC4517198/).","questions":[{"text":"Does partner sperm activate development, contribute inherited nuclear DNA, or have a variable fate?","id":"input-fate-q01","kind":"definition"},{"text":"Which breeding, cytological or parentage observations demonstrate that mechanism?","id":"input-fate-q02","kind":"provenance"}]}]}]},{"id":"genomic-continuity","name":"Genomic continuity","description":"Track ancestry, chromosome complements and transmission across generations.","layers":[{"id":"ancestry-and-cytotypes","name":"Ancestry and cytotypes","description":"Characterise parental contributions and chromosome-set variation.","findings":[{"id":"genomic-composition","name":"Genomic composition","description":"Record evidence for ancestral genome contributions and measured ploidy, retaining uncertainty where markers cannot resolve ancestry.","questions":[{"text":"Which ancestral taxa are supported by nuclear and mitochondrial evidence, and where do those signals disagree?","id":"genomic-composition-q01","kind":"provenance"},{"text":"Which ploidy levels occur in the sampled populations, and how were they measured?","id":"genomic-composition-q02","kind":"measurement"}]}]},{"id":"generational-transmission","name":"Generational transmission","description":"Connect somatic genome composition to gametes and offspring.","findings":[{"id":"retention-and-exclusion","name":"Retention and exclusion","description":"Record which genomes pass between generations and any demonstrated elimination or replacement. Water-frog research illustrates why retained ancestry and current partner contributions require separate records. Source: [Scientific Reports study](https://www.nature.com/articles/s41598-017-12942-y).","questions":[{"text":"Which genome components are retained, excluded or newly incorporated during gametogenesis and fertilisation?","id":"retention-and-exclusion-q01","kind":"measurement"},{"text":"Does transmission evidence support a continuing lineage or repeated formation of new hybrids?","id":"retention-and-exclusion-q02","kind":"boundary"}]}]}]},{"id":"population-persistence","name":"Population persistence","description":"Evaluate local reproductive systems and the consequences of changing their participants.","layers":[{"id":"local-system","name":"Local reproductive system","description":"Describe the partners, cytotypes and recruitment observed together at a site.","findings":[{"id":"population-specific-dependence","name":"Population-specific dependence","description":"Attach reproductive conclusions to sampled populations and dates. European water-frog research reports geographic variation in genome exclusion and reproductive systems. Source: [Distribution and evolution of western European water frogs](https://pmc.ncbi.nlm.nih.gov/articles/PMC12462689/).","questions":[{"text":"Which klepton cytotypes and compatible partner taxa coexist during the breeding period?","id":"population-specific-dependence-q01","kind":"measurement"},{"text":"What evidence demonstrates successful recruitment under the local reproductive arrangement?","id":"population-specific-dependence-q02","kind":"provenance"}]}]},{"id":"intervention-assessment","name":"Intervention assessment","description":"Evaluate proposed changes against the demonstrated reproductive system.","findings":[{"id":"dependency-sensitive-actions","name":"Dependency-sensitive actions","description":"Assess removal, introduction, captive breeding or translocation using partner compatibility and genome transmission evidence; record unknown outcomes explicitly.","questions":[{"text":"Would the proposed action remove a required partner or create contact with a new compatible taxon?","id":"dependency-sensitive-actions-q01","kind":"action"},{"text":"Which reproductive outcomes must be resolved before deciding whether the proposed action is appropriate?","id":"dependency-sensitive-actions-q02","kind":"action"}]}]}]},{"id":"evidence-and-external-alignment","name":"Evidence and external alignment","description":"What the world already says about this thing, gathered so the model can be checked against it.","layers":[{"id":"reported-evidence","name":"Reported evidence","description":"Findings from the breadth pass, kept separate from the structural claims.","findings":[{"id":"evidence-confidence-notes","name":"Check these first","description":"Recalled without web access and unsourced; every item is a lead to verify.","questions":[{"text":"Which of these check these first hold for the sense of klepton this model covers, and on what evidence?","id":"evidence-confidence-notes-q01","kind":"provenance"}]},{"id":"evidence-identifiers","name":"Identifiers and schemes","description":"Recalled without web access and unsourced; every item is a lead to verify.","questions":[{"text":"Which of these identifiers and schemes hold for the sense of klepton this model covers, and on what evidence?","id":"evidence-identifiers-q01","kind":"provenance"}]},{"id":"evidence-real-world-use","name":"Real-world use","description":"Recalled without web access and unsourced; every item is a lead to verify.","questions":[{"text":"Which of these real-world use hold for the sense of klepton this model covers, and on what evidence?","id":"evidence-real-world-use-q01","kind":"provenance"}]},{"id":"evidence-neighbours","name":"Neighbouring kinds and how to tell them apart","description":"Recalled without web access and unsourced; every item is a lead to verify.","questions":[{"text":"Which of these neighbouring kinds and how to tell them apart hold for the sense of klepton this model covers, and on what evidence?","id":"evidence-neighbours-q01","kind":"provenance"}]}]}]}]},"agentConduct":{"may":["Explain what a klepton is in biology and give documented examples from reliable sources.","Use the kl. notation correctly when recording hybrid-derived animals in data."],"mustNot":["Treat a klepton as a human organisation or social group.","Present a klepton as a normal species in conservation or taxonomic records.","Collect or move animals of klepton populations, such as water frogs, without permission from wildlife authorities.","Release hybrid animals into wild populations of their parent species."],"requiresHuman":["Decisions on conservation status or management of hybrid-dependent populations."]},"ethics":{"considerations":["Hybrid-dependent populations can affect the survival of the species they depend on.","Misclassification can lead to wrong conservation decisions."],"affectedParties":["Wild animal populations","Conservation managers","Researchers"]},"owners":{"steward":"Nobody owns the concept; wildlife authorities answer for wild populations.","roles":[],"masterSystems":["Biodiversity occurrence and taxonomic databases"]},"relations":[{"target":"Q16521","type":"parent","note":"registry parent class"}],"interaction":{"identity":{"applicability":"required","items":["Vercy registry: vr.tr.klepton","Wikidata: Q931051 (https://www.wikidata.org/wiki/Q931051)"]},"properties":{"applicability":"required","items":["Taxonomic treatment: klepton designation, alternative treatment, disputed, unresolved; qualified by authority and date","Reproductive partner dependence: Partner taxon linked to dependent population, reproductive role and evidence","Reproductive mechanism: hybridogenesis, gynogenesis, other documented mechanism, mixed, unresolved","Genome transmission pattern: Genome retained, excluded, recombined or incorporated; qualified by reproductive stage and population","Ploidy: Number of chromosome sets, expressed as multiples of x","Local reproductive continuity: recruitment demonstrated, recruitment unverified, recruitment failure observed, unresolved","Partner availability: Compatible partners confirmed, absent in surveyed sample, seasonal overlap missing, unresolved"]},"recognition":{"applicability":"not-applicable","items":[]},"capabilities":{"applicability":"required","items":["Assess whether a proposed klepton designation is supported by reproductive and genetic evidence.","Map dependent populations to their demonstrated reproductive partners.","Compare local genome transmission systems without assuming uniform behaviour across the taxon.","Identify observations needed to distinguish persistent lineages from transient hybrids.","Flag conservation or translocation proposals that could disrupt or create reproductive dependencies."]},"hazards":{"applicability":"required","items":["Moving hybrid animals can spread them into new areas and displace parent species.","Naming confusion can hide the real status of a population."]},"interfaces":{"applicability":"required","items":["The International Code of Zoological Nomenclature, issued by the International Commission on Zoological Nomenclature, governs applicable zoological names but does not establish klepton as a formal taxonomic rank."]},"context":{"applicability":"required","items":["The klepton concept is used in zoology to describe hybrid lineages whose reproduction depends on a related species.","It is encountered in research on hybridisation, reproductive systems and conservation where preserving a lineage may also require preserving its reproductive partner."]}},"sources":[{"title":"Wikidata item Q931051: klepton","url":"https://www.wikidata.org/wiki/Q931051","note":"identity and sense of the item"},{"title":"Wikipedia: Klepton","url":"https://en.wikipedia.org/wiki/Klepton","note":"general description of the item"}],"openQuestions":["Does the underlying registry source confirm the biological meaning of klepton, and why was it assigned to SOC / SOC.ORG?","Which authoritative definition should govern inclusion of hybridogenetic, gynogenetic and other reproductively dependent entities?","How should populations with different degrees of reproductive independence affect the circumscription of a named klepton?","What evidence threshold distinguishes a persistent klepton lineage from recurrent hybrid formation?","Does an existing Vercy world model already cover this concept, requiring a registry link rather than a separate publication?"],"resources":{"spec":"/models/things/publications/thing-q931051/spec.json"},"provenance":{"origin":"thing registry research (pass 2)","builtFrom":["models/things/publications/thing-q931051/spec.json"],"providers":["Codex"],"researchStatus":"unreviewed","generatedAt":"2026-09-12T10:46:36Z","builder":"tools/build_cards.py@1.0.0"},"completeness":{"sections":{"classifiers":"filled","whatItIs":"filled","purpose":"filled","distinguishingFeatures":"filled","structure":"filled","agentConduct":"filled","ethics":"filled","owners":"filled","relations":"filled","interaction.identity":"filled","interaction.properties":"filled","interaction.recognition":"not-applicable","interaction.capabilities":"filled","interaction.hazards":"filled","interaction.interfaces":"filled","interaction.context":"filled","sources":"filled"},"notes":{"_gate":"Published through the thing publication gate 1.0 on 2026-10-07T10:10:33+00:00: completeness 1.00; sense: Wikidata description; 2 live sources; review by codex: pass","interaction.recognition":"A klepton is a biological grouping defined by ancestry and reproductive dependence, which cannot be established from visual appearance alone."},"score":1.0}}