{
    "model": {
        "rank": 5273,
        "code": "thing-q154626",
        "model_id": "vr.tr.cytoskeleton",
        "name": "cytoskeleton",
        "purpose": "Let an agent explain the cytoskeleton and its components, relay their structure, dynamics and functions from cell biology sources, describe its role in disease and drug action, and distinguish its subsystems.",
        "family": "Thing Registry",
        "category": "Physical world and living systems",
        "status": "research-draft",
        "kind": "thing",
        "plane": "PHY",
        "domain": "PHY.LIV",
        "industry": "",
        "version": "",
        "url": "/models/thing/q154626/",
        "tier": 2,
        "score": 68,
        "payload": {
            "layer": "wikidata",
            "aliases": [
                "actin cytoskeleton",
                "intermediate filament cytoskeleton",
                "cortical cytoskeleton",
                "microtubule cytoskeleton",
                "spectrin-associated cytoskeleton",
                "perinuclear theca",
                "postsynaptic cytoskeleton",
                "presynaptic cytoskeleton",
                "septin cytoskeleton",
                "Prokaryotic cytoskeleton",
                "cortical actin cytoskeleton",
                "postsynaptic actin cytoskeleton",
                "presynaptic actin cytoskeleton"
            ],
            "aliasCount": 13,
            "merged": 17,
            "knownIn": 68,
            "facets": null,
            "markers": [],
            "lexicalClass": "",
            "senseRank": null,
            "alsoRegisteredAs": "vr.tr.cytoskeleton",
            "source": {
                "dataset": "wikidata",
                "item": "Q154626",
                "url": "https://www.wikidata.org/wiki/Q154626",
                "license": "CC0 1.0"
            }
        },
        "research": {
            "vercy": "1.0-draft",
            "publication": {
                "status": "research-draft",
                "adjudicationStatus": "unreviewed",
                "publishableCanonical": false,
                "generatedAt": "2026-09-12T22:24:12Z",
                "providers": [
                    "Claude"
                ],
                "breadth": "written by Claude from model knowledge without web access - no source was read, every claim is a lead to verify",
                "pass": 2,
                "wave": 3,
                "engine": "claude"
            },
            "metaModel": {
                "id": "THING-Q154626",
                "registryId": "vr.tr.cytoskeleton",
                "name": "cytoskeleton",
                "version": "0.2.0-wave.3",
                "entryKind": "thing",
                "family": "Thing Registry",
                "domain": [
                    "PHY.LIV"
                ],
                "status": "research-draft"
            },
            "canonicalUrl": "https://ver.cy/models/thing/q154626/",
            "model": {
                "registry_id": "vr.tr.cytoskeleton",
                "name": "cytoskeleton",
                "purpose": "Let an agent explain the cytoskeleton and its components, relay their structure, dynamics and functions from cell biology sources, describe its role in disease and drug action, and distinguish its subsystems.",
                "definition": "The network of protein filaments in the cytoplasm of cells that gives them shape, mechanical strength and internal organisation and drives movement, made up in eukaryotes of actin microfilaments, intermediate filaments and microtubules with associated motor and cross-linking proteins, and in bacteria and archaea of related proteins; the cytoskeleton underlies cell division, transport, muscle contraction and migration.",
                "what_it_is_for": "Shaping, organising and moving cells.",
                "affordances": [
                    "explain components and dynamics",
                    "relay functions in division, transport and movement",
                    "describe roles in disease and drugs",
                    "distinguish subsystems"
                ],
                "distinguishing_features": [
                    "Three filament systems in eukaryotes",
                    "Dynamic assembly and disassembly",
                    "Motor proteins",
                    "Prokaryotic homologues"
                ],
                "appearance": "Not visible to the eye; seen by microscopy as networks of filaments stained in fluorescent colours.",
                "visual_identification": [
                    "Filament network in the cytoplasm",
                    "Actin, intermediate filaments and microtubules",
                    "The extracellular matrix is outside the cell; the cell wall is a separate structure"
                ],
                "physical_properties": [
                    {
                        "quantity": "actin filament diameter",
                        "typical_range": "about 7",
                        "unit": "nm",
                        "note": ""
                    },
                    {
                        "quantity": "intermediate filament diameter",
                        "typical_range": "about 10",
                        "unit": "nm",
                        "note": ""
                    },
                    {
                        "quantity": "microtubule diameter",
                        "typical_range": "about 25",
                        "unit": "nm",
                        "note": ""
                    }
                ],
                "families_and_kinds": [
                    "actin cytoskeleton including the cortical cytoskeleton",
                    "intermediate filament cytoskeleton",
                    "microtubule cytoskeleton",
                    "spectrin-associated membrane skeleton",
                    "specialised structures such as the sperm perinuclear theca",
                    "prokaryotic cytoskeleton"
                ],
                "related_models": [
                    {
                        "relation": "is a kind of",
                        "target": "non-membrane-bounded organelle",
                        "why": "in registry terms"
                    },
                    {
                        "relation": "includes",
                        "target": "microtubule",
                        "why": "a filament system"
                    },
                    {
                        "relation": "includes",
                        "target": "actin filament",
                        "why": "a filament system"
                    },
                    {
                        "relation": "drives",
                        "target": "cell division",
                        "why": "through the spindle and contractile ring"
                    }
                ],
                "identifiers": [
                    {
                        "scheme": "Gene Ontology",
                        "value_or_pattern": "GO:0005856",
                        "note": "cytoskeleton"
                    }
                ],
                "standards_and_regulation": [
                    "No regulation; research standards and nomenclature"
                ],
                "failure_modes_and_hazards": [
                    "Diseases from cytoskeletal mutations such as some cardiomyopathies and neuropathies",
                    "Drug targets with side effects, such as microtubule inhibitors",
                    "Confusing subsystems"
                ],
                "in_scope": [],
                "out_of_scope": [],
                "characteristics": []
            },
            "sources": [],
            "structure": {
                "bundles": [
                    {
                        "id": "structure",
                        "name": "Structure",
                        "description": "Components.",
                        "rationale": "Science.",
                        "layers": [
                            {
                                "id": "filaments",
                                "name": "Filaments",
                                "description": "The three systems.",
                                "findings": [
                                    {
                                        "id": "filaments-finding",
                                        "name": "Filaments",
                                        "description": "Filaments.",
                                        "questions": [
                                            {
                                                "text": "What are actin filaments, intermediate filaments and microtubules, and how do they differ?",
                                                "kind": "definition"
                                            },
                                            {
                                                "text": "Is the structure cytoskeletal or extracellular?",
                                                "kind": "boundary"
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "proteins",
                                "name": "Proteins",
                                "description": "Associated proteins.",
                                "findings": [
                                    {
                                        "id": "proteins-finding",
                                        "name": "Proteins",
                                        "description": "Proteins.",
                                        "questions": [
                                            {
                                                "text": "What motor, nucleating and cross-linking proteins organise the cytoskeleton?",
                                                "kind": "definition"
                                            },
                                            {
                                                "text": "Which entry fits the specific protein?",
                                                "kind": "action"
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "function",
                        "name": "Function",
                        "description": "Functions.",
                        "rationale": "Science.",
                        "layers": [
                            {
                                "id": "dynamics",
                                "name": "Dynamics",
                                "description": "Dynamics.",
                                "findings": [
                                    {
                                        "id": "dynamics-finding",
                                        "name": "Dynamics",
                                        "description": "Dynamics.",
                                        "questions": [
                                            {
                                                "text": "How do filaments assemble, disassemble and generate force?",
                                                "kind": "provenance"
                                            },
                                            {
                                                "text": "Which references are standard?",
                                                "kind": "provenance"
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "roles",
                                "name": "Roles",
                                "description": "Roles in the cell.",
                                "findings": [
                                    {
                                        "id": "roles-finding",
                                        "name": "Roles",
                                        "description": "Roles.",
                                        "questions": [
                                            {
                                                "text": "How does the cytoskeleton drive division, transport, migration and muscle contraction?",
                                                "kind": "provenance"
                                            },
                                            {
                                                "text": "Which entry fits cell motility?",
                                                "kind": "action"
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "medicine",
                        "name": "Medicine",
                        "description": "Disease and drugs.",
                        "rationale": "Application.",
                        "layers": [
                            {
                                "id": "disease",
                                "name": "Disease",
                                "description": "Disease.",
                                "findings": [
                                    {
                                        "id": "disease-finding",
                                        "name": "Disease",
                                        "description": "Disease.",
                                        "questions": [
                                            {
                                                "text": "Which diseases involve cytoskeletal defects, with findings attributed?",
                                                "kind": "provenance"
                                            },
                                            {
                                                "text": "Is the user asking about a personal condition, which needs a clinician?",
                                                "kind": "boundary"
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "drugs",
                                "name": "Drugs",
                                "description": "Drug targets.",
                                "findings": [
                                    {
                                        "id": "drugs-finding",
                                        "name": "Drugs",
                                        "description": "Drugs.",
                                        "questions": [
                                            {
                                                "text": "How do drugs such as taxanes and vinca alkaloids act on the cytoskeleton?",
                                                "kind": "provenance"
                                            },
                                            {
                                                "text": "Which entry fits microtubule inhibitor?",
                                                "kind": "action"
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "context",
                        "name": "Context",
                        "description": "Evolution and study.",
                        "rationale": "Study.",
                        "layers": [
                            {
                                "id": "evolution",
                                "name": "Evolution",
                                "description": "Evolution.",
                                "findings": [
                                    {
                                        "id": "evolution-finding",
                                        "name": "Evolution",
                                        "description": "Evolution.",
                                        "questions": [
                                            {
                                                "text": "How did the cytoskeleton evolve, and what are its prokaryotic homologues?",
                                                "kind": "provenance"
                                            },
                                            {
                                                "text": "Which entry fits prokaryotic cytoskeleton?",
                                                "kind": "action"
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "methods",
                                "name": "Methods",
                                "description": "Methods.",
                                "findings": [
                                    {
                                        "id": "methods-finding",
                                        "name": "Methods",
                                        "description": "Methods.",
                                        "questions": [
                                            {
                                                "text": "How is the cytoskeleton studied, from fluorescence microscopy to cryo-electron tomography?",
                                                "kind": "provenance"
                                            },
                                            {
                                                "text": "Which sources are cited?",
                                                "kind": "provenance"
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    }
                ]
            },
            "openQuestions": [
                "Should each filament system be a separate entry?",
                "How should Gene Ontology terms be linked?",
                "The registry entry has merged aliases naming subsystems; should they be split off?"
            ],
            "statistics": {
                "bundles": 4,
                "layers": 8,
                "findings": 8,
                "questions": 16
            }
        },
        "draft": {
            "generator": "vr.draft.v3",
            "status": "draft-generated",
            "researched": false,
            "archetype": "organism",
            "method": "Written from the archetype playbook - what this kind of thing needs beyond identity and provenance - and from the structure that recurred across 6,333 models already researched by two engines. Applied to this entry by rule. No source was read for this thing and no claim here is researched. This entry carries no facets of its own, so they were inferred from its domain - a guess about a whole domain applied to one thing.",
            "facetsInferred": true,
            "nextPass": "A researcher replaces this draft with a sourced specification. Treat every sentence below as a proposal to argue with.",
            "purpose": "Give an agent a durable, checkable way to recognise a cytoskeleton, record what state it is in, and decide what may be done with it.",
            "whatItIs": "Let an agent explain the cytoskeleton and its components, relay their structure, dynamics and functions from cell biology sources, describe its role in disease and drug action, and distinguish its subsystems.",
            "characteristics": {
                "substance": "living",
                "origin": "natural",
                "agency": "autonomous",
                "mobility": "self-moving"
            },
            "whatYouCanDoWithIt": [
                "observed and measured"
            ],
            "distinguishingFeatures": [
                "Names folded into this entry, which a task may need to split apart again: actin cytoskeleton, intermediate filament cytoskeleton, cortical cytoskeleton, microtubule cytoskeleton, spectrin-associated cytoskeleton, perinuclear theca, postsynaptic cytoskeleton, presynaptic cytoskeleton, septin cytoskeleton, Prokaryotic cytoskeleton, cortical actin cytoskeleton, postsynaptic actin cytoskeleton.",
                "17 finer distinctions are held as aliases rather than separate entries, because telling them apart needs a task that asks for it.",
                "Described in 68 Wikipedia languages, which is a measure of how widely the thing is known, not of how important it is."
            ],
            "openQuestionsForResearch": [
                "Which of the bundles below does a real task actually need, and which are ceremony?",
                "What does this thing have that the facets do not capture at all?",
                "Which neighbouring kind is most often confused with a cytoskeleton, and on what evidence are they told apart?"
            ],
            "whatItIsMadeOf": "a living thing",
            "physicalCharacter": [
                "Moves under its own power or of its own accord.",
                "Acts on its own behalf.",
                "These come from the domain this entry sits in rather than from the entry itself, so treat them as a first guess about the whole domain applied to one thing."
            ],
            "whatCanBeDoneWithIt": [
                "observe it, measure it, record its state"
            ],
            "howItIsRecognised": [
                "Recognised by diagnostic characters, and those differ by life stage - the young of many kinds are mistaken for other kinds entirely. A sighting is settled by characters, a photograph and a place, not by a name alone."
            ],
            "relatedModels": [
                {
                    "relation": "covers",
                    "note": "Finer kinds folded into this entry because telling them apart needs a task that asks for it. Each is a model waiting to be split out when one does.",
                    "targets": [
                        "actin cytoskeleton",
                        "intermediate filament cytoskeleton",
                        "cortical cytoskeleton",
                        "microtubule cytoskeleton",
                        "spectrin-associated cytoskeleton",
                        "perinuclear theca",
                        "postsynaptic cytoskeleton",
                        "presynaptic cytoskeleton",
                        "septin cytoskeleton",
                        "Prokaryotic cytoskeleton",
                        "cortical actin cytoskeleton",
                        "postsynaptic actin cytoskeleton"
                    ]
                }
            ],
            "standing": "Described in 68 Wikipedia languages, which measures how widely it is written about rather than how important or how common it is. 17 finer distinctions are held inside this entry as names rather than as separate models.",
            "structure": {
                "bundles": [
                    {
                        "id": "identity-and-classification",
                        "name": "Identity, naming and classification",
                        "description": "How an agent tells one cytoskeleton from another, and a cytoskeleton from things that resemble it.",
                        "rationale": "Recognition comes before every other claim. Without stable identity nothing else in the model can be trusted to be about the same thing twice.",
                        "layers": [
                            {
                                "id": "naming-and-identifiers",
                                "name": "Names and identifiers",
                                "description": "The names this thing goes by and the identifiers that survive translation and time.",
                                "findings": [
                                    {
                                        "id": "preferred-name-and-aliases",
                                        "name": "Preferred name, aliases and local names",
                                        "description": "Which name to use, which names mean the same thing, and which merely sound similar.",
                                        "questions": [
                                            {
                                                "id": "preferred-name-and-aliases-q01",
                                                "text": "What identifies and describes the name of a cytoskeleton, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "preferred-name-and-aliases-q02",
                                                "text": "Who or what asserted this about the name of a cytoskeleton, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "preferred-name-and-aliases-q03",
                                                "text": "What may an agent decide or do once the name of a cytoskeleton is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    },
                                    {
                                        "id": "stable-identifiers",
                                        "name": "Stable identifiers and external keys",
                                        "description": "Identifiers that keep pointing at this kind of thing across systems and languages.",
                                        "questions": [
                                            {
                                                "id": "stable-identifiers-q01",
                                                "text": "What identifies and describes an identifier for a cytoskeleton, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "stable-identifiers-q02",
                                                "text": "Who or what asserted this about an identifier for a cytoskeleton, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "stable-identifiers-q03",
                                                "text": "What may an agent decide or do once an identifier for a cytoskeleton is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "classification-and-granularity",
                                "name": "Classification and granularity",
                                "description": "Where a cytoskeleton sits among kinds, and how finely a task needs to cut it.",
                                "findings": [
                                    {
                                        "id": "kind-and-parents",
                                        "name": "Kind, parents and neighbouring kinds",
                                        "description": "The classes this thing belongs to and the ones it is next to.",
                                        "questions": [
                                            {
                                                "id": "kind-and-parents-q01",
                                                "text": "What identifies and describes the kind of a cytoskeleton, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "kind-and-parents-q02",
                                                "text": "Who or what asserted this about the kind of a cytoskeleton, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "kind-and-parents-q03",
                                                "text": "What may an agent decide or do once the kind of a cytoskeleton is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    },
                                    {
                                        "id": "distinguishing-features",
                                        "name": "Distinguishing features",
                                        "description": "What separates a cytoskeleton from the things most often confused with it.",
                                        "questions": [
                                            {
                                                "id": "distinguishing-features-q01",
                                                "text": "What identifies and describes what distinguishes a cytoskeleton, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "distinguishing-features-q02",
                                                "text": "Who or what asserted this about what distinguishes a cytoskeleton, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "distinguishing-features-q03",
                                                "text": "What may an agent decide or do once what distinguishes a cytoskeleton is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "form-and-composition",
                        "name": "Form, composition and condition",
                        "description": "What a cytoskeleton is made of, what shape and size it takes, and what state it is in.",
                        "rationale": "A physical thing can be measured, and a model that cannot record its measurements cannot support any decision that depends on them.",
                        "layers": [
                            {
                                "id": "form-and-dimensions",
                                "name": "Form and dimensions",
                                "description": "Shape, size, mass and the ranges these take across examples.",
                                "findings": [
                                    {
                                        "id": "measurable-dimensions",
                                        "name": "Measurable dimensions and their units",
                                        "description": "The quantities worth recording and the units they are recorded in.",
                                        "questions": [
                                            {
                                                "id": "measurable-dimensions-q01",
                                                "text": "What identifies and describes the dimensions of a cytoskeleton, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "measurable-dimensions-q02",
                                                "text": "Who or what asserted this about the dimensions of a cytoskeleton, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "measurable-dimensions-q03",
                                                "text": "What may an agent decide or do once the dimensions of a cytoskeleton is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "material-and-composition",
                                "name": "Material and composition",
                                "description": "Anatomy, parts and life stage.",
                                "findings": [
                                    {
                                        "id": "parts-and-materials",
                                        "name": "Parts and life stages",
                                        "description": "The parts of a cytoskeleton and the stages it passes through.",
                                        "questions": [
                                            {
                                                "id": "parts-and-materials-q01",
                                                "text": "What identifies and describes the composition of a cytoskeleton, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "parts-and-materials-q02",
                                                "text": "Who or what asserted this about the composition of a cytoskeleton, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "parts-and-materials-q03",
                                                "text": "What may an agent decide or do once the composition of a cytoskeleton is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "condition-and-integrity",
                                "name": "Condition and integrity",
                                "description": "Wear, damage, health and the difference between working and broken.",
                                "findings": [
                                    {
                                        "id": "condition-assessment",
                                        "name": "Condition, damage and health",
                                        "description": "How the condition of a cytoskeleton is judged and by whom.",
                                        "questions": [
                                            {
                                                "id": "condition-assessment-q01",
                                                "text": "What identifies and describes the condition of a cytoskeleton, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "condition-assessment-q02",
                                                "text": "Who or what asserted this about the condition of a cytoskeleton, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "condition-assessment-q03",
                                                "text": "What may an agent decide or do once the condition of a cytoskeleton is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "place-and-movement",
                        "name": "Place and movement",
                        "description": "Where a cytoskeleton is, how it got there and where it may go (self-moving).",
                        "rationale": "A thing that can move needs its position recorded with a time, or every later claim about it is about a place it has left.",
                        "layers": [
                            {
                                "id": "location-and-placement",
                                "name": "Location and placement",
                                "description": "Position, containment and the reference frame the position is given in.",
                                "findings": [
                                    {
                                        "id": "position-and-frame",
                                        "name": "Position, container and reference frame",
                                        "description": "Where a cytoskeleton is, and what that position is measured against.",
                                        "questions": [
                                            {
                                                "id": "position-and-frame-q01",
                                                "text": "What identifies and describes the location of a cytoskeleton, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "position-and-frame-q02",
                                                "text": "Who or what asserted this about the location of a cytoskeleton, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "position-and-frame-q03",
                                                "text": "What may an agent decide or do once the location of a cytoskeleton is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "movement-and-transfer",
                                "name": "Movement and transfer",
                                "description": "How a cytoskeleton moves on its own.",
                                "findings": [
                                    {
                                        "id": "movement-events",
                                        "name": "Movement events and constraints",
                                        "description": "What counts as a movement, what records it and what limits it.",
                                        "questions": [
                                            {
                                                "id": "movement-events-q01",
                                                "text": "What identifies and describes the movement of a cytoskeleton, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "movement-events-q02",
                                                "text": "Who or what asserted this about the movement of a cytoskeleton, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "movement-events-q03",
                                                "text": "What may an agent decide or do once the movement of a cytoskeleton is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "state-and-lifecycle",
                        "name": "State and lifecycle",
                        "description": "The states a cytoskeleton passes through and the events that move it between them.",
                        "rationale": "Most decisions about a thing depend on what state it is in now, which is a claim with a time on it, not a property.",
                        "layers": [
                            {
                                "id": "lifecycle-stages",
                                "name": "Lifecycle stages",
                                "description": "From coming into existence to ceasing to be one of these.",
                                "findings": [
                                    {
                                        "id": "stages-and-transitions",
                                        "name": "Stages and transitions",
                                        "description": "The stages worth naming and what moves a cytoskeleton between them.",
                                        "questions": [
                                            {
                                                "id": "stages-and-transitions-q01",
                                                "text": "What identifies and describes the lifecycle of a cytoskeleton, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "stages-and-transitions-q02",
                                                "text": "Who or what asserted this about the lifecycle of a cytoskeleton, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "stages-and-transitions-q03",
                                                "text": "What may an agent decide or do once the lifecycle of a cytoskeleton is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "observations-and-status",
                                "name": "Observations and current status",
                                "description": "What is observed about a cytoskeleton, how often and by whom.",
                                "findings": [
                                    {
                                        "id": "observation-record",
                                        "name": "Observation record",
                                        "description": "How an observation of a cytoskeleton is recorded so that it can be superseded rather than overwritten.",
                                        "questions": [
                                            {
                                                "id": "observation-record-q01",
                                                "text": "What identifies and describes an observation of a cytoskeleton, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "observation-record-q02",
                                                "text": "Who or what asserted this about an observation of a cytoskeleton, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "observation-record-q03",
                                                "text": "What may an agent decide or do once an observation of a cytoskeleton is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "taxonomic-placement",
                        "name": "Taxonomic placement and identification",
                        "description": "What cytoskeleton is called scientifically, by whose authority, and how a specimen is recognised.",
                        "rationale": "Names change with revisions while the organism does not, so a name without its authority and date cannot be reconciled later.",
                        "layers": [
                            {
                                "id": "accepted-name",
                                "name": "Accepted name and synonymy",
                                "description": "The current binomial, its author and date, and the names it replaced.",
                                "findings": [
                                    {
                                        "id": "name-and-authority",
                                        "name": "Accepted name, authority and synonyms",
                                        "description": "What to record so a later revision can be applied without losing the old data.",
                                        "questions": [
                                            {
                                                "id": "name-and-authority-q01",
                                                "text": "What is the accepted scientific name of cytoskeleton, with its author and year, and which synonyms does it absorb?",
                                                "kind": "definition"
                                            },
                                            {
                                                "id": "name-and-authority-q02",
                                                "text": "Which taxonomic authority is being followed, and what would a competing authority say?",
                                                "kind": "provenance"
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "field-identification",
                                "name": "Field identification",
                                "description": "The characters that separate cytoskeleton from what it is confused with.",
                                "findings": [
                                    {
                                        "id": "diagnostic-characters",
                                        "name": "Diagnostic characters and confusable kinds",
                                        "description": "Features that settle an identification, and those that only suggest it.",
                                        "questions": [
                                            {
                                                "id": "diagnostic-characters-q01",
                                                "text": "Which observable characters distinguish cytoskeleton from its closest look-alikes, and at which life stage are they visible?",
                                                "kind": "boundary"
                                            },
                                            {
                                                "id": "diagnostic-characters-q02",
                                                "text": "What evidence is enough to record an identification as confirmed rather than provisional?",
                                                "kind": "provenance"
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "life-cycle-and-condition",
                        "name": "Life stages, condition and behaviour",
                        "description": "What cytoskeleton does over its life and how its state is judged.",
                        "rationale": "Stage and condition change what may be done - handled, treated, harvested, left alone - and both are observations with a time.",
                        "layers": [
                            {
                                "id": "stages-and-reproduction",
                                "name": "Stages and reproduction",
                                "description": "Development from origin to death, including forms that look like different animals.",
                                "findings": [
                                    {
                                        "id": "life-stages",
                                        "name": "Life stages and their marks",
                                        "description": "The stages worth recording and what tells them apart.",
                                        "questions": [
                                            {
                                                "id": "life-stages-q01",
                                                "text": "What stages does cytoskeleton pass through, and what observable change marks each transition?",
                                                "kind": "definition"
                                            },
                                            {
                                                "id": "life-stages-q02",
                                                "text": "Which stages are so unlike the adult that they are mistaken for another kind?",
                                                "kind": "boundary"
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "health-and-condition",
                                "name": "Health and condition",
                                "description": "How condition is assessed and by whom.",
                                "findings": [
                                    {
                                        "id": "condition-assessment",
                                        "name": "Condition assessment",
                                        "description": "Signs of health, injury and stress, and who is competent to judge them.",
                                        "questions": [
                                            {
                                                "id": "condition-assessment-q01",
                                                "text": "How is the condition of cytoskeleton judged, on what scale, and by whom?",
                                                "kind": "measurement"
                                            },
                                            {
                                                "id": "condition-assessment-q02",
                                                "text": "What condition obliges an agent to act, and what act is permitted to whom?",
                                                "kind": "action"
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "range-and-status",
                        "name": "Habitat, range and conservation status",
                        "description": "Where cytoskeleton lives and how threatened it is.",
                        "rationale": "Range and status govern what is lawful, and both are assessments with an assessor and a year rather than facts of nature.",
                        "layers": [
                            {
                                "id": "habitat-and-range",
                                "name": "Habitat and range",
                                "description": "Native range, introductions and the habitat it needs.",
                                "findings": [
                                    {
                                        "id": "range-record",
                                        "name": "Range, introductions and habitat",
                                        "description": "What to record about where it is found and how it got there.",
                                        "questions": [
                                            {
                                                "id": "range-record-q01",
                                                "text": "What is the native range of cytoskeleton, where has it been introduced, and what habitat does it require?",
                                                "kind": "definition"
                                            },
                                            {
                                                "id": "range-record-q02",
                                                "text": "What distinguishes a native record from an introduced or vagrant one?",
                                                "kind": "boundary"
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "protection-status",
                                "name": "Protection and legal status",
                                "description": "Conservation assessments and the rules that follow from them.",
                                "findings": [
                                    {
                                        "id": "status-and-rules",
                                        "name": "Conservation status and legal consequence",
                                        "description": "The assessment, its date, and what it forbids or requires where.",
                                        "questions": [
                                            {
                                                "id": "status-and-rules-q01",
                                                "text": "What conservation status does cytoskeleton hold, under which assessment and year?",
                                                "kind": "provenance"
                                            },
                                            {
                                                "id": "status-and-rules-q02",
                                                "text": "What does that status forbid or require of an agent, and in which jurisdiction?",
                                                "kind": "action"
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "provenance-and-evidence",
                        "name": "Provenance, evidence and time",
                        "description": "Where every claim about a cytoskeleton came from and when it held.",
                        "rationale": "A claim without a source and a time cannot be superseded, only overwritten, and an agent that overwrites loses the ability to explain itself.",
                        "layers": [
                            {
                                "id": "source-and-authority",
                                "name": "Source and authority",
                                "description": "Who said it, on what evidence, and how strongly.",
                                "findings": [
                                    {
                                        "id": "claim-provenance",
                                        "name": "Claim provenance and confidence",
                                        "description": "The authority behind each claim about a cytoskeleton and how confident it is.",
                                        "questions": [
                                            {
                                                "id": "claim-provenance-q01",
                                                "text": "What identifies and describes a claim about a cytoskeleton, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "claim-provenance-q02",
                                                "text": "Who or what asserted this about a claim about a cytoskeleton, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "claim-provenance-q03",
                                                "text": "What may an agent decide or do once a claim about a cytoskeleton is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "time-and-versions",
                                "name": "Time, versions and supersession",
                                "description": "When a claim was true, when it was learnt, and what replaced it.",
                                "findings": [
                                    {
                                        "id": "validity-and-supersession",
                                        "name": "Validity period and supersession",
                                        "description": "How an old claim about a cytoskeleton is retired without being erased.",
                                        "questions": [
                                            {
                                                "id": "validity-and-supersession-q01",
                                                "text": "What identifies and describes the validity of a claim about a cytoskeleton, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "validity-and-supersession-q02",
                                                "text": "Who or what asserted this about the validity of a claim about a cytoskeleton, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "validity-and-supersession-q03",
                                                "text": "What may an agent decide or do once the validity of a claim about a cytoskeleton is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    }
                ]
            },
            "statistics": {
                "bundles": 8,
                "layers": 17,
                "findings": 19,
                "questions": 51
            }
        }
    }
}