{
    "model": {
        "rank": 7901,
        "code": "thing-q189933",
        "model_id": "vr.tr.microtubule",
        "name": "microtubule",
        "purpose": "Let an agent explain microtubules, relay structure, dynamics, functions and medical relevance from cell biology sources, describe the named microtubule populations, and distinguish microtubules from actin filaments, intermediate filaments, centrioles as structures built from microtubules and bacterial tubulin homologues.",
        "family": "Thing Registry",
        "category": "Physical world and living systems",
        "status": "research-draft",
        "kind": "thing",
        "plane": "PHY",
        "domain": "PHY.LIV",
        "industry": "",
        "version": "",
        "url": "/models/thing/q189933/",
        "tier": 2,
        "score": 53,
        "payload": {
            "layer": "wikidata",
            "aliases": [
                "spindle microtubule",
                "cytoplasmic microtubule",
                "nuclear microtubule",
                "intraconoid microtubule",
                "dendritic microtubule",
                "subpellicular microtubule",
                "post-anaphase array microtubule",
                "neurotubule",
                "neuropil thread"
            ],
            "aliasCount": 9,
            "merged": 9,
            "knownIn": 53,
            "facets": null,
            "markers": [],
            "lexicalClass": "",
            "senseRank": null,
            "alsoRegisteredAs": "vr.tr.microtubule",
            "source": {
                "dataset": "wikidata",
                "item": "Q189933",
                "url": "https://www.wikidata.org/wiki/Q189933",
                "license": "CC0 1.0"
            }
        },
        "research": {
            "vercy": "1.0-draft",
            "publication": {
                "status": "research-draft",
                "adjudicationStatus": "unreviewed",
                "publishableCanonical": false,
                "generatedAt": "2026-09-13T21:11:32Z",
                "providers": [
                    "Claude"
                ],
                "breadth": "written by Claude from model knowledge without web access - no source was read, every claim is a lead to verify",
                "pass": 2,
                "wave": 4,
                "engine": "claude"
            },
            "metaModel": {
                "id": "THING-Q189933",
                "registryId": "vr.tr.microtubule",
                "name": "microtubule",
                "version": "0.2.0-wave.4",
                "entryKind": "thing",
                "family": "Thing Registry",
                "domain": [
                    "PHY.LIV"
                ],
                "status": "research-draft"
            },
            "canonicalUrl": "https://ver.cy/models/thing/q189933/",
            "model": {
                "registry_id": "vr.tr.microtubule",
                "name": "microtubule",
                "purpose": "Let an agent explain microtubules, relay structure, dynamics, functions and medical relevance from cell biology sources, describe the named microtubule populations, and distinguish microtubules from actin filaments, intermediate filaments, centrioles as structures built from microtubules and bacterial tubulin homologues.",
                "definition": "A hollow cylindrical polymer of alpha- and beta-tubulin, about 25 nanometres in diameter, forming part of the cytoskeleton of eukaryotic cells, with dynamic instability of growth and shrinkage described by Mitchison and Kirschner in 1984, organising the mitotic spindle, cilia and flagella and intracellular transport by motor proteins, found in populations the registry aliases name such as spindle, cytoplasmic, nuclear and dendritic microtubules, and specialised arrays such as subpellicular microtubules in trypanosomes and apicomplexans and intraconoid microtubules in the conoid of apicomplexan parasites; drugs such as taxanes and vinca alkaloids target microtubules in cancer therapy.",
                "what_it_is_for": "Cell shape, division, transport and motility.",
                "affordances": [
                    "explain structure and dynamics",
                    "relay functions",
                    "describe named populations",
                    "relay drug targeting"
                ],
                "distinguishing_features": [
                    "Hollow tube",
                    "Polarity",
                    "Dynamic instability",
                    "Drug target"
                ],
                "appearance": "Not visible to the eye; seen in fluorescence and electron microscopy as fine tubes.",
                "visual_identification": [
                    "Tubulin polymer tube of the cytoskeleton",
                    "Spindle, cytoplasmic, nuclear, intraconoid, dendritic and subpellicular microtubules",
                    "Actin filaments are thinner; intermediate filaments are rope-like; centrioles are built from microtubules; FtsZ is a bacterial homologue"
                ],
                "physical_properties": [
                    {
                        "quantity": "outer diameter",
                        "typical_range": "about 25",
                        "unit": "nm",
                        "note": ""
                    },
                    {
                        "quantity": "protofilaments",
                        "typical_range": "typically 13",
                        "unit": "count",
                        "note": ""
                    },
                    {
                        "quantity": "dynamic instability described",
                        "typical_range": "1984",
                        "unit": "year",
                        "note": "Mitchison and Kirschner"
                    }
                ],
                "families_and_kinds": [
                    "spindle microtubules",
                    "cytoplasmic microtubules",
                    "axonemal microtubules in cilia",
                    "neuronal and dendritic microtubules",
                    "parasite subpellicular and conoid microtubules"
                ],
                "related_models": [
                    {
                        "relation": "is a kind of",
                        "target": "cytoskeleton",
                        "why": "component"
                    },
                    {
                        "relation": "is made of",
                        "target": "tubulin",
                        "why": ""
                    },
                    {
                        "relation": "forms",
                        "target": "mitotic spindle",
                        "why": ""
                    },
                    {
                        "relation": "is targeted by",
                        "target": "paclitaxel",
                        "why": ""
                    }
                ],
                "identifiers": [],
                "standards_and_regulation": [
                    "Gene Ontology cellular component terms"
                ],
                "failure_modes_and_hazards": [
                    "Oversimplifying dynamics",
                    "Confusing cytoskeletal filament types",
                    "Overstating speculative roles such as in consciousness"
                ],
                "in_scope": [],
                "out_of_scope": [],
                "characteristics": []
            },
            "sources": [],
            "structure": {
                "bundles": [
                    {
                        "id": "understand",
                        "name": "Understand",
                        "description": "What a microtubule is.",
                        "rationale": "Science.",
                        "layers": [
                            {
                                "id": "definition",
                                "name": "Definition",
                                "description": "Definition.",
                                "findings": [
                                    {
                                        "id": "definition-finding",
                                        "name": "Definition",
                                        "description": "Definition.",
                                        "questions": [
                                            {
                                                "text": "What is a microtubule, and how does it differ from actin and intermediate filaments, centrioles and FtsZ?",
                                                "kind": "definition"
                                            },
                                            {
                                                "text": "Is the question about cell biology, cancer drugs, neurons or parasites?",
                                                "kind": "boundary"
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "populations",
                                "name": "Populations",
                                "description": "Named populations.",
                                "findings": [
                                    {
                                        "id": "populations-finding",
                                        "name": "Populations",
                                        "description": "Populations.",
                                        "questions": [
                                            {
                                                "text": "What are spindle, cytoplasmic, nuclear, dendritic, subpellicular and intraconoid microtubules?",
                                                "kind": "definition"
                                            },
                                            {
                                                "text": "Which entry fits the specific population?",
                                                "kind": "action"
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "biology",
                        "name": "Biology",
                        "description": "Structure and dynamics.",
                        "rationale": "Science.",
                        "layers": [
                            {
                                "id": "structure",
                                "name": "Structure",
                                "description": "Structure.",
                                "findings": [
                                    {
                                        "id": "structure-finding",
                                        "name": "Structure",
                                        "description": "Structure.",
                                        "questions": [
                                            {
                                                "text": "How do tubulin dimers assemble into protofilaments and tubes?",
                                                "kind": "provenance"
                                            },
                                            {
                                                "text": "Which references are standard?",
                                                "kind": "provenance"
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "dynamics",
                                "name": "Dynamics",
                                "description": "Dynamic instability.",
                                "findings": [
                                    {
                                        "id": "dynamics-finding",
                                        "name": "Dynamics",
                                        "description": "Dynamics.",
                                        "questions": [
                                            {
                                                "text": "What is dynamic instability, and how is it regulated?",
                                                "kind": "provenance"
                                            },
                                            {
                                                "text": "Which sources are cited?",
                                                "kind": "provenance"
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "function",
                        "name": "Function",
                        "description": "Functions.",
                        "rationale": "Application.",
                        "layers": [
                            {
                                "id": "division",
                                "name": "Division",
                                "description": "Cell division.",
                                "findings": [
                                    {
                                        "id": "division-finding",
                                        "name": "Division",
                                        "description": "Division.",
                                        "questions": [
                                            {
                                                "text": "How do spindle microtubules separate chromosomes?",
                                                "kind": "provenance"
                                            },
                                            {
                                                "text": "Which entry fits mitosis?",
                                                "kind": "action"
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "transport",
                                "name": "Transport",
                                "description": "Motor transport.",
                                "findings": [
                                    {
                                        "id": "transport-finding",
                                        "name": "Transport",
                                        "description": "Transport.",
                                        "questions": [
                                            {
                                                "text": "How do kinesin and dynein move cargo along microtubules?",
                                                "kind": "provenance"
                                            },
                                            {
                                                "text": "Which entry fits kinesin?",
                                                "kind": "action"
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "context",
                        "name": "Context",
                        "description": "Medicine.",
                        "rationale": "Context.",
                        "layers": [
                            {
                                "id": "drugs",
                                "name": "Drugs",
                                "description": "Cancer drugs.",
                                "findings": [
                                    {
                                        "id": "drugs-finding",
                                        "name": "Drugs",
                                        "description": "Drugs.",
                                        "questions": [
                                            {
                                                "text": "How do taxanes and vinca alkaloids affect microtubules?",
                                                "kind": "provenance"
                                            },
                                            {
                                                "text": "Which entry fits vinca alkaloid?",
                                                "kind": "action"
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "disease",
                                "name": "Disease",
                                "description": "Disease links.",
                                "findings": [
                                    {
                                        "id": "disease-finding",
                                        "name": "Disease",
                                        "description": "Disease.",
                                        "questions": [
                                            {
                                                "text": "How are microtubule-associated proteins such as tau linked to neurodegeneration?",
                                                "kind": "provenance"
                                            },
                                            {
                                                "text": "Which entry fits tau protein?",
                                                "kind": "action"
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    }
                ]
            },
            "openQuestions": [
                "Should parasite microtubule arrays be separate entries?",
                "How should Gene Ontology be linked?",
                "How should speculative hypotheses be flagged?"
            ],
            "statistics": {
                "bundles": 4,
                "layers": 8,
                "findings": 8,
                "questions": 16
            }
        },
        "draft": {
            "generator": "vr.draft.v3",
            "status": "draft-generated",
            "researched": false,
            "archetype": "organism",
            "method": "Written from the archetype playbook - what this kind of thing needs beyond identity and provenance - and from the structure that recurred across 6,333 models already researched by two engines. Applied to this entry by rule. No source was read for this thing and no claim here is researched. This entry carries no facets of its own, so they were inferred from its domain - a guess about a whole domain applied to one thing.",
            "facetsInferred": true,
            "nextPass": "A researcher replaces this draft with a sourced specification. Treat every sentence below as a proposal to argue with.",
            "purpose": "Give an agent a durable, checkable way to recognise a microtubule, record what state it is in, and decide what may be done with it.",
            "whatItIs": "Let an agent explain microtubules, relay structure, dynamics, functions and medical relevance from cell biology sources, describe the named microtubule populations, and distinguish microtubules from actin filaments, intermediate filaments, centrioles as structures built from microtubules and bacterial tubulin homologues.",
            "characteristics": {
                "substance": "living",
                "origin": "natural",
                "agency": "autonomous",
                "mobility": "self-moving"
            },
            "whatYouCanDoWithIt": [
                "observed and measured"
            ],
            "distinguishingFeatures": [
                "Names folded into this entry, which a task may need to split apart again: spindle microtubule, cytoplasmic microtubule, nuclear microtubule, intraconoid microtubule, dendritic microtubule, subpellicular microtubule, post-anaphase array microtubule, neurotubule, neuropil thread.",
                "9 finer distinctions are held as aliases rather than separate entries, because telling them apart needs a task that asks for it.",
                "Described in 53 Wikipedia languages, which is a measure of how widely the thing is known, not of how important it is."
            ],
            "openQuestionsForResearch": [
                "Which of the bundles below does a real task actually need, and which are ceremony?",
                "What does this thing have that the facets do not capture at all?",
                "Which neighbouring kind is most often confused with a microtubule, and on what evidence are they told apart?"
            ],
            "whatItIsMadeOf": "a living thing",
            "physicalCharacter": [
                "Moves under its own power or of its own accord.",
                "Acts on its own behalf.",
                "These come from the domain this entry sits in rather than from the entry itself, so treat them as a first guess about the whole domain applied to one thing."
            ],
            "whatCanBeDoneWithIt": [
                "observe it, measure it, record its state"
            ],
            "howItIsRecognised": [
                "Recognised by diagnostic characters, and those differ by life stage - the young of many kinds are mistaken for other kinds entirely. A sighting is settled by characters, a photograph and a place, not by a name alone."
            ],
            "relatedModels": [
                {
                    "relation": "covers",
                    "note": "Finer kinds folded into this entry because telling them apart needs a task that asks for it. Each is a model waiting to be split out when one does.",
                    "targets": [
                        "spindle microtubule",
                        "cytoplasmic microtubule",
                        "nuclear microtubule",
                        "intraconoid microtubule",
                        "dendritic microtubule",
                        "subpellicular microtubule",
                        "post-anaphase array microtubule",
                        "neurotubule",
                        "neuropil thread"
                    ]
                }
            ],
            "standing": "Described in 53 Wikipedia languages, which measures how widely it is written about rather than how important or how common it is. 9 finer distinctions are held inside this entry as names rather than as separate models.",
            "structure": {
                "bundles": [
                    {
                        "id": "identity-and-classification",
                        "name": "Identity, naming and classification",
                        "description": "How an agent tells one microtubule from another, and a microtubule from things that resemble it.",
                        "rationale": "Recognition comes before every other claim. Without stable identity nothing else in the model can be trusted to be about the same thing twice.",
                        "layers": [
                            {
                                "id": "naming-and-identifiers",
                                "name": "Names and identifiers",
                                "description": "The names this thing goes by and the identifiers that survive translation and time.",
                                "findings": [
                                    {
                                        "id": "preferred-name-and-aliases",
                                        "name": "Preferred name, aliases and local names",
                                        "description": "Which name to use, which names mean the same thing, and which merely sound similar.",
                                        "questions": [
                                            {
                                                "id": "preferred-name-and-aliases-q01",
                                                "text": "What identifies and describes the name of a microtubule, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "preferred-name-and-aliases-q02",
                                                "text": "Who or what asserted this about the name of a microtubule, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "preferred-name-and-aliases-q03",
                                                "text": "What may an agent decide or do once the name of a microtubule is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    },
                                    {
                                        "id": "stable-identifiers",
                                        "name": "Stable identifiers and external keys",
                                        "description": "Identifiers that keep pointing at this kind of thing across systems and languages.",
                                        "questions": [
                                            {
                                                "id": "stable-identifiers-q01",
                                                "text": "What identifies and describes an identifier for a microtubule, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "stable-identifiers-q02",
                                                "text": "Who or what asserted this about an identifier for a microtubule, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "stable-identifiers-q03",
                                                "text": "What may an agent decide or do once an identifier for a microtubule is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "classification-and-granularity",
                                "name": "Classification and granularity",
                                "description": "Where a microtubule sits among kinds, and how finely a task needs to cut it.",
                                "findings": [
                                    {
                                        "id": "kind-and-parents",
                                        "name": "Kind, parents and neighbouring kinds",
                                        "description": "The classes this thing belongs to and the ones it is next to.",
                                        "questions": [
                                            {
                                                "id": "kind-and-parents-q01",
                                                "text": "What identifies and describes the kind of a microtubule, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "kind-and-parents-q02",
                                                "text": "Who or what asserted this about the kind of a microtubule, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "kind-and-parents-q03",
                                                "text": "What may an agent decide or do once the kind of a microtubule is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    },
                                    {
                                        "id": "distinguishing-features",
                                        "name": "Distinguishing features",
                                        "description": "What separates a microtubule from the things most often confused with it.",
                                        "questions": [
                                            {
                                                "id": "distinguishing-features-q01",
                                                "text": "What identifies and describes what distinguishes a microtubule, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "distinguishing-features-q02",
                                                "text": "Who or what asserted this about what distinguishes a microtubule, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "distinguishing-features-q03",
                                                "text": "What may an agent decide or do once what distinguishes a microtubule is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "form-and-composition",
                        "name": "Form, composition and condition",
                        "description": "What a microtubule is made of, what shape and size it takes, and what state it is in.",
                        "rationale": "A physical thing can be measured, and a model that cannot record its measurements cannot support any decision that depends on them.",
                        "layers": [
                            {
                                "id": "form-and-dimensions",
                                "name": "Form and dimensions",
                                "description": "Shape, size, mass and the ranges these take across examples.",
                                "findings": [
                                    {
                                        "id": "measurable-dimensions",
                                        "name": "Measurable dimensions and their units",
                                        "description": "The quantities worth recording and the units they are recorded in.",
                                        "questions": [
                                            {
                                                "id": "measurable-dimensions-q01",
                                                "text": "What identifies and describes the dimensions of a microtubule, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "measurable-dimensions-q02",
                                                "text": "Who or what asserted this about the dimensions of a microtubule, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "measurable-dimensions-q03",
                                                "text": "What may an agent decide or do once the dimensions of a microtubule is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "material-and-composition",
                                "name": "Material and composition",
                                "description": "Anatomy, parts and life stage.",
                                "findings": [
                                    {
                                        "id": "parts-and-materials",
                                        "name": "Parts and life stages",
                                        "description": "The parts of a microtubule and the stages it passes through.",
                                        "questions": [
                                            {
                                                "id": "parts-and-materials-q01",
                                                "text": "What identifies and describes the composition of a microtubule, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "parts-and-materials-q02",
                                                "text": "Who or what asserted this about the composition of a microtubule, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "parts-and-materials-q03",
                                                "text": "What may an agent decide or do once the composition of a microtubule is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "condition-and-integrity",
                                "name": "Condition and integrity",
                                "description": "Wear, damage, health and the difference between working and broken.",
                                "findings": [
                                    {
                                        "id": "condition-assessment",
                                        "name": "Condition, damage and health",
                                        "description": "How the condition of a microtubule is judged and by whom.",
                                        "questions": [
                                            {
                                                "id": "condition-assessment-q01",
                                                "text": "What identifies and describes the condition of a microtubule, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "condition-assessment-q02",
                                                "text": "Who or what asserted this about the condition of a microtubule, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "condition-assessment-q03",
                                                "text": "What may an agent decide or do once the condition of a microtubule is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "place-and-movement",
                        "name": "Place and movement",
                        "description": "Where a microtubule is, how it got there and where it may go (self-moving).",
                        "rationale": "A thing that can move needs its position recorded with a time, or every later claim about it is about a place it has left.",
                        "layers": [
                            {
                                "id": "location-and-placement",
                                "name": "Location and placement",
                                "description": "Position, containment and the reference frame the position is given in.",
                                "findings": [
                                    {
                                        "id": "position-and-frame",
                                        "name": "Position, container and reference frame",
                                        "description": "Where a microtubule is, and what that position is measured against.",
                                        "questions": [
                                            {
                                                "id": "position-and-frame-q01",
                                                "text": "What identifies and describes the location of a microtubule, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "position-and-frame-q02",
                                                "text": "Who or what asserted this about the location of a microtubule, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "position-and-frame-q03",
                                                "text": "What may an agent decide or do once the location of a microtubule is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "movement-and-transfer",
                                "name": "Movement and transfer",
                                "description": "How a microtubule moves on its own.",
                                "findings": [
                                    {
                                        "id": "movement-events",
                                        "name": "Movement events and constraints",
                                        "description": "What counts as a movement, what records it and what limits it.",
                                        "questions": [
                                            {
                                                "id": "movement-events-q01",
                                                "text": "What identifies and describes the movement of a microtubule, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "movement-events-q02",
                                                "text": "Who or what asserted this about the movement of a microtubule, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "movement-events-q03",
                                                "text": "What may an agent decide or do once the movement of a microtubule is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "state-and-lifecycle",
                        "name": "State and lifecycle",
                        "description": "The states a microtubule passes through and the events that move it between them.",
                        "rationale": "Most decisions about a thing depend on what state it is in now, which is a claim with a time on it, not a property.",
                        "layers": [
                            {
                                "id": "lifecycle-stages",
                                "name": "Lifecycle stages",
                                "description": "From coming into existence to ceasing to be one of these.",
                                "findings": [
                                    {
                                        "id": "stages-and-transitions",
                                        "name": "Stages and transitions",
                                        "description": "The stages worth naming and what moves a microtubule between them.",
                                        "questions": [
                                            {
                                                "id": "stages-and-transitions-q01",
                                                "text": "What identifies and describes the lifecycle of a microtubule, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "stages-and-transitions-q02",
                                                "text": "Who or what asserted this about the lifecycle of a microtubule, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "stages-and-transitions-q03",
                                                "text": "What may an agent decide or do once the lifecycle of a microtubule is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "observations-and-status",
                                "name": "Observations and current status",
                                "description": "What is observed about a microtubule, how often and by whom.",
                                "findings": [
                                    {
                                        "id": "observation-record",
                                        "name": "Observation record",
                                        "description": "How an observation of a microtubule is recorded so that it can be superseded rather than overwritten.",
                                        "questions": [
                                            {
                                                "id": "observation-record-q01",
                                                "text": "What identifies and describes an observation of a microtubule, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "observation-record-q02",
                                                "text": "Who or what asserted this about an observation of a microtubule, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "observation-record-q03",
                                                "text": "What may an agent decide or do once an observation of a microtubule is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "taxonomic-placement",
                        "name": "Taxonomic placement and identification",
                        "description": "What microtubule is called scientifically, by whose authority, and how a specimen is recognised.",
                        "rationale": "Names change with revisions while the organism does not, so a name without its authority and date cannot be reconciled later.",
                        "layers": [
                            {
                                "id": "accepted-name",
                                "name": "Accepted name and synonymy",
                                "description": "The current binomial, its author and date, and the names it replaced.",
                                "findings": [
                                    {
                                        "id": "name-and-authority",
                                        "name": "Accepted name, authority and synonyms",
                                        "description": "What to record so a later revision can be applied without losing the old data.",
                                        "questions": [
                                            {
                                                "id": "name-and-authority-q01",
                                                "text": "What is the accepted scientific name of microtubule, with its author and year, and which synonyms does it absorb?",
                                                "kind": "definition"
                                            },
                                            {
                                                "id": "name-and-authority-q02",
                                                "text": "Which taxonomic authority is being followed, and what would a competing authority say?",
                                                "kind": "provenance"
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "field-identification",
                                "name": "Field identification",
                                "description": "The characters that separate microtubule from what it is confused with.",
                                "findings": [
                                    {
                                        "id": "diagnostic-characters",
                                        "name": "Diagnostic characters and confusable kinds",
                                        "description": "Features that settle an identification, and those that only suggest it.",
                                        "questions": [
                                            {
                                                "id": "diagnostic-characters-q01",
                                                "text": "Which observable characters distinguish microtubule from its closest look-alikes, and at which life stage are they visible?",
                                                "kind": "boundary"
                                            },
                                            {
                                                "id": "diagnostic-characters-q02",
                                                "text": "What evidence is enough to record an identification as confirmed rather than provisional?",
                                                "kind": "provenance"
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "life-cycle-and-condition",
                        "name": "Life stages, condition and behaviour",
                        "description": "What microtubule does over its life and how its state is judged.",
                        "rationale": "Stage and condition change what may be done - handled, treated, harvested, left alone - and both are observations with a time.",
                        "layers": [
                            {
                                "id": "stages-and-reproduction",
                                "name": "Stages and reproduction",
                                "description": "Development from origin to death, including forms that look like different animals.",
                                "findings": [
                                    {
                                        "id": "life-stages",
                                        "name": "Life stages and their marks",
                                        "description": "The stages worth recording and what tells them apart.",
                                        "questions": [
                                            {
                                                "id": "life-stages-q01",
                                                "text": "What stages does microtubule pass through, and what observable change marks each transition?",
                                                "kind": "definition"
                                            },
                                            {
                                                "id": "life-stages-q02",
                                                "text": "Which stages are so unlike the adult that they are mistaken for another kind?",
                                                "kind": "boundary"
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "health-and-condition",
                                "name": "Health and condition",
                                "description": "How condition is assessed and by whom.",
                                "findings": [
                                    {
                                        "id": "condition-assessment",
                                        "name": "Condition assessment",
                                        "description": "Signs of health, injury and stress, and who is competent to judge them.",
                                        "questions": [
                                            {
                                                "id": "condition-assessment-q01",
                                                "text": "How is the condition of microtubule judged, on what scale, and by whom?",
                                                "kind": "measurement"
                                            },
                                            {
                                                "id": "condition-assessment-q02",
                                                "text": "What condition obliges an agent to act, and what act is permitted to whom?",
                                                "kind": "action"
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "range-and-status",
                        "name": "Habitat, range and conservation status",
                        "description": "Where microtubule lives and how threatened it is.",
                        "rationale": "Range and status govern what is lawful, and both are assessments with an assessor and a year rather than facts of nature.",
                        "layers": [
                            {
                                "id": "habitat-and-range",
                                "name": "Habitat and range",
                                "description": "Native range, introductions and the habitat it needs.",
                                "findings": [
                                    {
                                        "id": "range-record",
                                        "name": "Range, introductions and habitat",
                                        "description": "What to record about where it is found and how it got there.",
                                        "questions": [
                                            {
                                                "id": "range-record-q01",
                                                "text": "What is the native range of microtubule, where has it been introduced, and what habitat does it require?",
                                                "kind": "definition"
                                            },
                                            {
                                                "id": "range-record-q02",
                                                "text": "What distinguishes a native record from an introduced or vagrant one?",
                                                "kind": "boundary"
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "protection-status",
                                "name": "Protection and legal status",
                                "description": "Conservation assessments and the rules that follow from them.",
                                "findings": [
                                    {
                                        "id": "status-and-rules",
                                        "name": "Conservation status and legal consequence",
                                        "description": "The assessment, its date, and what it forbids or requires where.",
                                        "questions": [
                                            {
                                                "id": "status-and-rules-q01",
                                                "text": "What conservation status does microtubule hold, under which assessment and year?",
                                                "kind": "provenance"
                                            },
                                            {
                                                "id": "status-and-rules-q02",
                                                "text": "What does that status forbid or require of an agent, and in which jurisdiction?",
                                                "kind": "action"
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    },
                    {
                        "id": "provenance-and-evidence",
                        "name": "Provenance, evidence and time",
                        "description": "Where every claim about a microtubule came from and when it held.",
                        "rationale": "A claim without a source and a time cannot be superseded, only overwritten, and an agent that overwrites loses the ability to explain itself.",
                        "layers": [
                            {
                                "id": "source-and-authority",
                                "name": "Source and authority",
                                "description": "Who said it, on what evidence, and how strongly.",
                                "findings": [
                                    {
                                        "id": "claim-provenance",
                                        "name": "Claim provenance and confidence",
                                        "description": "The authority behind each claim about a microtubule and how confident it is.",
                                        "questions": [
                                            {
                                                "id": "claim-provenance-q01",
                                                "text": "What identifies and describes a claim about a microtubule, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "claim-provenance-q02",
                                                "text": "Who or what asserted this about a claim about a microtubule, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "claim-provenance-q03",
                                                "text": "What may an agent decide or do once a claim about a microtubule is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            },
                            {
                                "id": "time-and-versions",
                                "name": "Time, versions and supersession",
                                "description": "When a claim was true, when it was learnt, and what replaced it.",
                                "findings": [
                                    {
                                        "id": "validity-and-supersession",
                                        "name": "Validity period and supersession",
                                        "description": "How an old claim about a microtubule is retired without being erased.",
                                        "questions": [
                                            {
                                                "id": "validity-and-supersession-q01",
                                                "text": "What identifies and describes the validity of a claim about a microtubule, and in what units or vocabulary?",
                                                "kind": "definition",
                                                "answer_data": [
                                                    "identifiers",
                                                    "types and classes",
                                                    "values with units",
                                                    "explicit unknowns"
                                                ]
                                            },
                                            {
                                                "id": "validity-and-supersession-q02",
                                                "text": "Who or what asserted this about the validity of a claim about a microtubule, by which method, and when was it true?",
                                                "kind": "provenance",
                                                "answer_data": [
                                                    "authority",
                                                    "method",
                                                    "evidence",
                                                    "event time",
                                                    "knowledge time"
                                                ]
                                            },
                                            {
                                                "id": "validity-and-supersession-q03",
                                                "text": "What may an agent decide or do once the validity of a claim about a microtubule is known, and what must it refuse?",
                                                "kind": "action",
                                                "answer_data": [
                                                    "permitted actions",
                                                    "preconditions",
                                                    "refusals",
                                                    "escalation"
                                                ]
                                            }
                                        ]
                                    }
                                ]
                            }
                        ]
                    }
                ]
            },
            "statistics": {
                "bundles": 8,
                "layers": 17,
                "findings": 19,
                "questions": 51
            }
        }
    }
}